Type
Subzone
Age range
22.96–22.44 Ma
Miocene
Aquitanian
G. dehiscens/P. kugleri CRSZ
P. kugleri LOSZ
P. pseudokugleri LOZ
22.9822.9622.9422.9222.9022.8822.8622.8422.8222.8022.7822.7622.7422.7222.7022.6822.6622.6422.6222.6022.5822.5622.5422.5222.5022.4822.4622.4422.42